Garden Park, Colorado, led to the discovery of a virtually complete specimen of Stegosaurus stenops (CARPENTER si: SMALL 1993, CARPENTER 1993, in press). The Small Stegosaurus Quarry has also produced a diverse vertebrate fauna, including many microvertebrates (Table 1). Except for the Stegosaurus specimen, all of the elements were disarticulated. Most of the vertebrate specimens were recovered from a lenticular gray lacustrine mudstone that is capped by a sequence of progressively thin sandstones separated by very thin mudstone. The sandstone becomes progressively thicker and vrider, and documents the growth of a crevasse splay complex (CARpENTER, in press). Fusian is abundant in the lacustrine mudstone, probably due to local fires accompanying a drought {CARFENTER in press}. Most of the fossils from the quarry have

A large pterodactyloid from the Morrison Formation 475 undergone diagenetic distortion. That some of these elements are ptero- saurian is certain as they posses extremely thin walls and, in some instances, internal struts (convergent with some birds}. Such struts are characteristic of later Cretaceous pterodactyloids, but are not restricted to those groups (WELLNHQFER 1991a). The pterosaur bones occur in light gray mudstones and greenish-gray fine-grained sandstones.

HowsE (1986), UNWIN (1992, 1995), PADIAN sc RAYNER (1993), and KELLNER {1995) have noted that the "Rhamphorhynchoidea" is a paraphyletic group; until the formal presentation of a new classificatory convention, we use the term "rhamphorhynchoid" in the informal, conventional sense {sensu UNwIN 1995), meaning those primitive ptero- saurs retaining long tails, wing finger phalanges chevron-shaped, not ovoid, in cross-section, large fifth pedal digits, occipital condyles facing posteriorly rather than posteroventrally, and separate nasal and antor- bital openings in the skull. Systematic paleontology

Pterosauria KAvP, 1834 Pterodactyloidea PLrENrNGER, 1901

Kepodactylus, n, g,

Etymology: Greek kepos meaning "garden, for Garden Park, Colorado, where the fossils were found; Greek dalctylos, meaning "finger". Type species: Kepodactylus insperatus n. sp. Type locality: DMNH Loc. 611, 1992 Stegosaurus stenops Quarry, near the top of the lower member of the Morrison Formation, Garden Park, Colorado {CARPENTER, in press).

Diagnosis: Large pterodactyloid having an estimated wingspan of 2.5 m. Cervical centrum elongated as in Pterodactylus but with postzygapophyses extended beyond centrum; cervical centrum lacking diapophyses of "rhamphor- hynchoids" and accessory exapophyses of azhdarchids; pneumatic toramen posteriorly located, elongated oval located on centrum rather than on neural arch as in "rhamphorhynchoids." First wing phalanx without longitudinal groove of "rhamphorhynchoids" and ovoid in cross-section, Deltopectoral crest large, rectangular, not hatchet-shaped as in "rhamphorhynchoids" and nyctosaurids, not warped as in pteranodontids.

Kepodactylus t'rtsperatus n. sp, Etymology: Latin insperatus, meaning "unexpected." Holotype: DMNH 21684, consisting of a cervical vertebra, a left humerus, the left proximal phalanx of the wing finger, the distal end of the right proximal wing finger phalanx, the proximal end of the left second wing finger phalanx, and a metatarsal. Type locality: As for genus. Diagnosis: As for genus. 476 Jerald D. Harris and Kenneth Carpenter 1 I I / I I I I I I B C Fig. 1. Cervical vertebra of Kepodactylus insperatus. A: Anterior view. B: Dorsal view. C: Ventral view. D: Right lateral view. Arrow indicates pneumatic foramen

Scale = 2 cm.